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Published Manta Research

Anderson RC, Adam MS, Goes J (2011) From monsoons to mantas: seasonal distribution of Manta alfredi in the Maldives. Fisheries Oceanography 20: 104-113

Anderson RC, Adam MS, Kitchen-Wheeler A, Stevens G (2010) Extent and economic value of manta ray watching in the Maldives. Tourism in Marine Environments 7: 15-27

Ari C (2011) Encephalization and brain organization of mobulid rays (Myliobatiformes, Elasmobranchii) with ecological perspectives. The Open Anatomy Journal 3: 1-13

Ari C (2014a) Long-term body pigmentation changes on a manta ray (Mobulidae). Biological Journal of the Linnean Society 

Ari C (2014b) Rapid coloration changes of manta rays (Mobulidae) Biological Journal of the Linnean Society: 180-193

Ari C, Correia JP (2008) Role of sensory cues on searching behavior of a captive Manta birostris (Chondrichtyes, Mobulidae). Zoo Biology 27: 294-304

Clark TB, Papastamatiou YP (2007) Intestinal eversion in a free-ranging manta ray. Coral Reefs 27

Couturier LIE, Dudgeon CL, Pollock KH, Jaine FRA, Bennett MB, Townsend KA, Weeks SJ, Richardson AJ (2014) Population dynamics of the reef manta ray Manta alfredi in eastern Australia. Coral Reefs 33: 329–342

Couturier LIE, Jaine FRA, Townsend KA, Weeks SJ, Richardson AJ, Bennett MB (2011) Distribution, site affinity and regional movements of the manta ray, Manta alfredi (Krefft, 1868), along the east coast of Australia. Marine and Freshwater Research 62: 628–637

Couturier LIE, Marshall AD, Jaine FRA, Kashiwagi T, Pierce SJ, Townsend KA, Weeks SJ, Bennett MB, Richardson AJ (2012) Biology, ecology and conservation of the Mobulidae. Journal of Fish Biology 80: 1075-1119

Deakos MH (2010) Paired-laser photogrammetry as a simple and accurate system for measuring the body size of free-ranging manta rays Manta alfredi. Aquatic Biology 10: 1-10

Deakos MH (2011) The reproductive ecology of resident manta rays (Manta alfredi) off Maui, Hawaii, with an emphasis on body size. Environmental Biology of Fishes 

Deakos MH, Baker JD, Bejder L (2011) Characteristics of a manta ray Manta alfredi population off Maui, Hawaii, and implications for management Marine Ecology-Progress Series 429: 234-260

Dewar H, Mous P, Domeier M, Muljadi A, Pet J, Whitty J (2008) Movements and site fidelity of the giant manta ray, Manta birostris, in the Komodo Marine Park, Indonesia. Marine Biology 155: 121-133

Freedman R, Shouraseni SR (2012) Spatial patterning of Manta birostris in United States east coast offshore habitat. Applied Geography 32: 652–659

Germanov ES, Marshall AD (2014) Running the gauntlet: regional movement patterns of Manta alfredi through a complex of parks and fisheries. PLoS ONE 9(10): e110071. doi:10.1371/journal.pone.0110071

Graham RT, Witt MJ, Castellanos DW, Remolina F, Maxwell S, Godley BJ, Hawkes LA (2012) Satellite tracking of manta rays highlights challenges to their conservation. PLoS ONE 7: e36734

Jaine FRA, Couturier LIE, Weeks SJ, Townsend KA, Bennett MB, Fiora K, Richardson AJ (2012) When giants turn up: sighting trends, environmental influences and habitat use of the manta ray Manta alfredi at a coral reef. PLoS ONE 7 e46170

Jaine FRA, Rohner CA, Weeks SJ, Couturier LIE, Bennett MB, Townsend KA, Richardson AJ (2014) Movements and habitat use of reef manta rays off eastern Australia: offshore excursions, deep diving and eddy affinity revealed by satellite telemetry. Marine Ecology-Progress Series 73-86

Kashiwagi T, Broderick D, Lance SL, Bennett MB, Ovenden J (2012a) Development and characterization of ten microsatellite loci for the reef manta ray Manta alfredi Conservation Genetics Resources 4: 1055-1058

Kashiwagi T, Marshall AD, Bennett MB, Ovenden JR (2011) Habitat segregation and mosaic sympatry of the two species of manta ray in the Indian and Pacific Oceans: Manta alfredi and M. birostris. Marine Biodiversity Records 4: 1-8

Kashiwagi T, Marshall AD, Bennett MB, Ovenden JR (2012b) The genetic signature of recent speciation in manta rays (Manta alfredi and M. birostris). Molecular Phylogenetics and Evolution 64: 212-218

Kitchen-Wheeler A-M (2013) The behaviour and ecology of Alfred mantas (Manta alfredi) in the Maldives. PhD Thesis, Newcastle University, UK

Kitchen-Wheeler A (2010) Visual identification of individual manta ray (Manta alfredi) in the Maldives Islands, Western Indian Ocean  Marine Biology Research 4: 351-363

Kitchen-Wheeler A, Ari C, Edwards AJ (2012) Population estimates of Alfred mantas (Manta alfredi) in central Maldives atolls: North Male, Ari and Baa. Environmental Biology of Fishes 93: 557-575

Luiz OJ, Balboni AP, Kodja G, Andrade M, Marum H (2009) Seasonal occurrences of Manta birostris (Chondricthyes: Mobulidae) in southeastern Brazil. Ichthyological Research 56: 96-99

Marshall AD, Bennett MB (2010a) The frequency and effect of shark-inflicted bite injuries to the reef manta ray Manta alfredi. African Journal of Marine Science 32: 573-580

Marshall AD, Bennett MB (2010b) Reproductive ecology of the reef manta ray Manta alfredi in southern Mozambique. Journal of Fish Biology 77: 169-190

Marshall AD, Compagno LJV, Bennett MB (2009) Redescription of the genus Manta with resurrection of Manta alfredi (Krefft,1868) (Chondrichthyes; Myliobatoidei; Mobulidae). Zootaxa 2301: 1-28

Marshall AD, Dudgeon CL, Bennett MB (2011) Size and structure of a photographically identified population of manta rays Manta alfredi in southern Mozambique. Marine Biology 158    1111-1124

Marshall AD, Pierce SJ (2012) The use and abuse of photographic identification in sharks and rays. Journal of Fish Biology 80: 1361–1379

Marshall AD, Pierce SJ, Bennett M (2008) Morphological measurements of manta rays (Manta birostris) with a description of a foetus from the east coast of Southern Africa. Zootaxa 1717: 24-30

Mourier J (2012) Manta rays in the Marquesas Islands: first records of Manta birostris in French Polynesia and most easterly location of Manta alfredi in the Pacific Ocean, with notes on their distribution Journal of Fish Biology 81: 2053-2058

O'Shea OR, Kingsford MJ, Seymour J (2010) Tide-related periodicity of manta rays and sharks to cleaning stations on a coral reef. Marine & Freshwater Research 61: 65-73

O’Malley MP, Lee-Brooks K, Medd HB (2013) The Global Economic Impact of Manta Ray Watching Tourism PLoS ONE

Papastamatiou YP, DeSalles PA, McCauley DJ (2012) Area-restricted searching by manta rays and their

response to spatial scale in lagoon habitats. Marine Ecology-Progress Series 456: 233-244

Tomita T, Toda M, Ueda K, Uchida S, Nakaya K (2012) Live-bearing manta ray: how the embryo acquires oxygen without placenta and umbilical cord. Biology Letters 8: 721-724 



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